As I understand it there's a significant pressured reservoir of water beneath the gorge, fed from multiple sources, which converge in such a way that they can produce different results at the same times from the outlets. Russet Well and Slop Moll seem to be quite intimately connected, but presumably complex overflow levels between them determine what happens where. They're fed by the Speedwell streamway and Lumbago Sump (at least),and possibly have some connection to Speedwell Pot. It's also possible sinking water in Cave Dale could resurge here, going under the Halfway House or the Swine Hole in a lower passage.
Also now Peakshole Sough has been shown to be a significant third resurgence from this system - not by draining mine workings, but via an upwelling in the floor of the sough, only 20 or so metres inside the level. In drought it's usually level with the floor, and any wetter it overflows into the passage and out into Peakshole Water. Dye traces showed this to be fed from surface sinking water in Cowlow Nick, the Son of Longcliffe sump and Longcliffe Mine (at least - I suspect Rowter Hole and Winnats Head could be added to the list). It hadn't really been used for detecting before that, so it may be significant. I think Jim L hoped to dive it but the slot is far too small to get into and I think he said a camera lowered down didn't look promising. We had a very strong trace here and a very weak one at RW, but at identical times, so it's clearly mixing with other water just after PS.
There's a large number of other potential inputs other than Peak-Speedwell, not least Treak Cliff and Blue John caverns, which both drain water via some unknown passages en-route - possibly some BJ water going into Odin Sough instead, but Treak Cliff is heading more toward Peak as I remember. I still think there's another streamway system north of, and roughly parallel to Speedwell, along the base of Long Cliff or under the road. There's a phreatic passage system under the shale at the far end of PS that Jim and Mich Ping dug out, that could well be an early higher-level part of this system, once connected to Goosehill Cave.