Longest one down south would almost certainly be the Blacknor / Sandy Hole system on Portland. The old separate surveys prior to the connection showed 600m+ for Blacknor Hole and c.400m for Sandy Hole. The connection did not add a great deal of passage, but the subsequent extension from the end of Ammonite Passage in Sandy Hole (1995) certainly did. The survey is yet to be published, but my recollection is that the Ammonite extensions are probably in excess of 2km (it seems a LOT more as there is nothing bigger than hands and knees, and most of it lower). I would therefore make an educated guess that the system is 3km+ in total length. Hope that's useful...